Jakob von Uexküll (1864–1944) was a biologist, but the impact of his work has been perhaps stronger and more persistent in philosophy and the humanities than in the natural sciences. As one of the contributors to this book observes, Uexküll's conception of biology is “more at home among the disciplines composing the Geisteswissenschaften [humanities] than those included in the Naturwissenschaften [sciences], insofar as Uexküll's biology put Verstehen [understanding] before Erklären [explaining].” Uexküll began his career as an experimentalist of a “proto-behaviorist” kind but came rapidly to be dissatisfied with the reduction of physiology to action-and-reaction chains. He argued that the animal-as-machine, the model that lies behind behaviorism, cannot be the starting point of biology. The relation between cause and effect in a machine is direct: the wings of a windmill, say, transfer their movement to a cog even if no one is there to perceive the transfer. In contrast, in...
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