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Ecosystems are popularly understood as having essential parts, species, which are necessary for the functioning of the whole. Early theorists supposed that ecological communities would predictably reemerge in the same form if destroyed, while later conservationists worried that they might fall apart with species extinctions. Evidence suggests that multispecies communities are constantly being transformed by chance encounters, historical accidents, and parasitic invasions. Unruly assemblages, novel ecosystems, are flourishing and proliferating in unexpected places. Transformative encounters, seductive moments that generate new entangled modes of coexistence, are constantly taking place as beings capture one another in reciprocal embraces. Diverse critters are ever ready to negotiate new symbiotic agreements.

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